Representing the state of the art in evolutionary paleobiology, this book provides a much-needed overview of this rapidly changing field. An influx of ideas and techniques both from other areas of biology and from within paleobiology itself have resulted in numerous recent advances, including increased recognition of the relationships between ecological and evolutionary theory, renewed vigor in the study of ecological communities over geologic timescales, increased understanding of biogeographical patterns, and new mathematical approaches to studying the form and structure of plants and animals.

Contributors to this volume—a veritable who's who of eminent researchers—present the results of original research and new theoretical developments, and provide directions for future studies. Individually wide ranging, these papers all share a debt to the work of James W. Valentine, one of the founders of modern evolutionary paleobiology. This volume's unified approach to the study of life on earth will be a major contribution to paleobiology, evolution, and ecology.

With all the recent advances in molecular and evolutionary biology, one could almost wonder why we need the fossil record. Molecular sequence data can resolve taxonomic relationships, experiments with fruit flies demonstrate evolution and development in real time, and field studies of Galapagos finches have provided the strongest evidence for natural selection ever measured in the wild. What, then, can fossils teach us that living organisms cannot?

Evolutionary Patterns demonstrates the rich variety of clues to evolution that can be gleaned from the fossil record. Chief among these are the major trends and anomalies in species development revealed only by "deep time," such as periodic mass extinctions and species that remain unchanged in form for millions of years. Contributors explore modes of development, the tempo of speciation and extinction, and macroevolutionary patterns and trends. The result is an important contribution to paleobiology and evolutionary biology, and a spirited defense of the fossil record as a crucial tool for understanding evolution and development.

The contributors are Ann F. Budd, Efstathia Bura, Leo W. Buss, Mike Foote, Jörn Geister, Stephen Jay Gould, Eckart Hâkansson, Jean-Georges Harmelin, Lee-Ann C. Hayek, Jeremy B. C. Jackson, Kenneth G. Johnson, Nancy Knowlton, Scott Lidgard, Frank K. McKinney, Daniel W. McShea, Ross H. Nehm, Beth Okamura, John M. Pandolfi, Paul D. Taylor, and Erik Thomsen.

Although the species is one of the fundamental units of biological classification, there is remarkably little consensus among biologists about what defines a species, even within distinct sub-disciplines. The literature of paleobiology, in particular, is littered with qualifiers and cautions about applying the term to the fossil record or equating such species with those recognized among living organisms. In Species and Speciation in the Fossil Record, experts in the field examine how they conceive of species of fossil animals and consider the implications these different approaches have for thinking about species in the context of macroevolution.

After outlining views of the Modern Synthesis of evolutionary disciplines and detailing the development within paleobiology of quantitative methods for documenting and analyzing variation within fossil assemblages, contributors explore the challenges of recognizing and defining species from fossil specimens—and offer potential solutions. Addressing both the tempo and mode of speciation over time, they show how with careful interpretation and a clear species concept, fossil species may be sufficiently robust for meaningful paleobiological analyses. Indeed, they demonstrate that the species concept, if more refined, could unearth a wealth of information about the interplay between species origins and extinctions, between local and global climate change, and greatly deepen our understanding of the evolution of life.